December 2019: Changes to evaluation system

Written by Ezequiel Nicolazzi , Paul VanRaden , George Wiggans

December 01, 2019

Updates to crossbred evaluations
By Ezequiel Nicolazzi (CDCB), George Wiggans (CDCB), Leigh Walton (CDCB) and Paul VanRaden (USDA-ARS-AGIL)

Two important updates will be implemented in the crossbred evaluations in December 2019:

Change in F1’s threshold for the breed of evaluation to be the breed of the ID: Since the introduction of crossbred evaluations in April 2019, Breed Ban Representation (BBR) is instrumental in determining the breed of evaluation of all animals. Until now, for F1 animals, the breed code of the animal ID deter the breed of evaluation even if the BBR value is only the second highest value if the 2 highest BBR breeds are in the range of 45-55% and the second breed than 10% from the first breed.

Effective with the December 2019 evaluation, the simplified rule will maintain the breed of the ID if its BBR is higher than 40%. In that case an animal with Hwill maintain the HO breed of evaluation if its HO BBR is higher than 40%. This change currently impacts less than 500 animals in the evaluation.

Change in calculation of reliabilities for crossbred animals: The calculations of reliability (REL) and the inbreeding of future progeny (EFI) rely on the relationship between the animal and the predictor population. Since crossbred animals do not have a predictor population for PTA (purebred SNP effects a blended in based on BBR values), their reliability and EFI estimates have previously been obtained using a multi-breed reference population. This strategy nearly 30% lower reliabilities for animals with BBR<90 compared to animals with BBR>90. A further concern was that crossbred animals with stronger links to the refere population (e.g. BBR close to the 90 threshold) receive similar reliabilities than animals having a mixed-breed genetic makeup (e.g. F1’s, or animals with contributions from more than 2 breeds).

Effective December 2019, following a full review of the methodology and results, the most connected purebred reference population of each crossbred anim (determined by the breed of evaluation used) will be used instead. In addition, a differential weighting of the traditional and genomic components will be applied, by giving more weight on the traditional component for animals with traditional reliability above 30% (animals with phenotypes)changes make it even more important to emphasize that genomic evaluations on crossbred animals are useful for animals being bred to animals of the same breed of evaluation (e.g. towards “purity”), whereas are not an advisable tool for animals in a rotational crossbreeding program.

As expected, no effect will be observed on purebred animals, whereas reliabilities and PTAs of crossbred animals will be impacted. (Note that reliabilities a to weight the traditional and genomic components of an evaluation.) The greatest changes will be observed on animals with BBR close to the 90% threshold, as their reliabilities will be the ones mostly changing (upwards) with this new strategy. Reliabilities will be more closely linked to distribution, as animals having
more purebred composition will receive a reliability and PTA estimates similar (with differences in weighting, obviously) to an animal. Considering that animals with BBR close to the 90% threshold have most of their SNP effects based on that same breed on which relatio are obtained from, this change makes reliability estimates more accurate.

Correction to use of foreign fertility evaluations
By Ezequiel Nicolazzi (CDCB), Leigh Walton (CDCB) and Paul VanRaden (USDA-ARS-AGIL)

When extending the MACE information to include Mastitis into the evaluation in August 2019, an incorrect trait order when comparing reliabilities caused M fertility evaluations to be used incorrectly for some bulls. This resulted in more differences between traditional and genomic evaluations than expected, which affected highly-reliable Holstein bulls and animals related to them. Test results on the error repair indicate the divergence between traditional and genomic these bulls should be resolved and is expected to result in slightly different genomic PTAs propagating throughout the population, due to better SNP estimation.

We thank Ryan Starkenburg (ABS) for identifying the misalignment and the industry review committees for providing feedback.

End of line policy in effect on all CDCB files
By Ezequiel Nicolazzi (CDCB)

For legacy purposes, CDCB had continued to change the EOL’s for zipped files to DOS EOL (CR LF) whereas unzipped files had LINUX EOL (LF). Since September 2019, all CDCB files (public and private) have been distributed using a standardized Linux “end of line” (LS EOL) control character. The December run will be the first evaluation release in which all files distributed by CDCB, irrespectively of being zipped or not, will have a standardized LINUX EOL (LF) character.

IP change in all our servers
By Ezequiel Nicolazzi (CDCB) and Frank Ross (CDCB)

In October, CDCB completed a transition to new IPs on all our servers. All users connecting to CDCB systems using the IP address (68.69.3.XXX) will not be able to access our systems. Please use:
 Web services: www.uscdcb.com / queries.uscdcb.com
 Redmine services (collaborators): redmine.uscdcb.com
 (s)FTP (public and private): ftp.uscdcb.com